Rebel forces claim to have taken control of the Syrian capital after storming through the country in less than two weeks.
According to reports, authoritarian ruler Bashar al Assad has fled Damascus on a plane headed to an unknown destination, ending his 24-year rule.
While Syrians celebrate in the streets with chants for freedom, attention will also turn to the insurgents and what happens next.
Who are the rebels?
The initial assault on regime forces, which began in the northern city of Aleppo last week, was carried out by a variety of Mr Assad’s opponents.
This included rebels under the banner of the Syrian National Army, backed by Turkey, but the offensive has mostly been led by jihadi group Hayat Tahrir al Sham (HTS).
Once known as the Nusra Front, a former wing of al Qaeda, HTS is said to have around 30,000 troops and has long been designated a terrorist group by the US and Russia.
It was formed to oppose the Syrian government and was founded by Abu Muhammed al Jolani, cutting ties with al Qaeda in 2016 and making an effort to appear moderate.
Image: Rebel fighters in the city of Homs. Pic: Reuters
As rebels entered Aleppo, video showed him issuing orders by phone, forbidding fighters from entering homes and reminding them to protect citizens.
Aron Lund, a fellow at think-tank Century International, said Mr Jolani and HTS have clearly changed, while adding they remain “pretty hardline”.
“It’s PR, but the fact they are engaging in this effort at all shows they are no longer as rigid as they once were,” he said.
“Old-school al Qaeda or the Islamic State would never have done that.”
But the US’s Commission on International Religious Freedom said in 2022 that despite the “rebrand”, HTS “restricts religious freedom” and threatens the safety of religious minorities.
What plans do they have?
Mr Jolani, himself designated a terrorist by the US in 2013, has tried to reassure Syrian minorities who fear jihadists.
In 2023, he allowed the first Christian mass in years in the northwestern city of Idlib, and on Wednesday insisted he would protect residents of a Christian town south of Aleppo.
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Syrian rebels announce Damascus ‘freed’
The group’s political plans remain to be seen, with Robert Ford, former US ambassador to Syria, telling Sky News he has not seen “detailed programs”.
“We don’t know exactly what they’re going to do,” he said. “I would say here that after a 13-year incredibly bloody and vicious civil war, right now it’s a day for Syrians to celebrate, and to hope for things that will turn out better.
“There will be plenty of hard work quickly waiting for them.”
Why has this happened now?
These anti-Assad victories are years in the making, more than a decade on from the start of Syrian civil war.
But the rapid rebel progress has stunned the international community.
Image: Key locations in Syria as well as the Iraqi town of Al Qaim, where troops sought refuge.
Former head of the British Army’s chemical weapons unit, Hamish De Bretton-Gordon, said the timing is “no coincidence”.
“With Hezbollah much diminished, the Iranian proxy in the region, and also other Iranian militias, it gave the rebels an opportunity,” he told Sky News.
“With Russia taking so many forces away from Syria to bolster their special military operation in Ukraine, it left Assad exposed and the rebels have really taken advantage of it.”
What other nations have interests in Syria?
The assault had raised the prospect of another front reopening in the Middle East, at a time when US-backed Israel has been fighting Hamas in Gaza and Hezbollah in Lebanon, both Iran-allied groups.
Russia, which was Mr Assad’s main international backer, is also preoccupied with its war in Ukraine.
Previous military intervention by Russia and Iran – alongside support from other groups – had allowed Mr Assad to remain in power and retain 70% of Syria under his control.
The US has about 900 troops in northeast Syria to guard against a resurgence by the Islamic State, which was defeated in Syria and Iraq by 2019, by a US-led coalition that included Kurdish fighters and UK air support.
Turkey, which also opposes Kurdish expansion, has forces in Syria alongside its reported influence with the broad alliance of opposition forces.
Iran’s response to Israeli attacks on its nuclear facilities is “self-defence” and a “matter of principle”, the Iranian ambassador to the UK has told Sky News.
Speaking exclusively to The World With Yalda Hakim, Seyed Ali Mousavi said the “barbaric Israeli regime” is “violating international law” – describing Israel’s actions in recent days as “an act of aggression against the Iranian people”.
The conflict between Israel and Iran – once played out in a series of proxy wars – has escalated in the past three days.
Image: Sky’s Yalda Hakim spoke to Iran’s ambassador to the UK, Seyed Ali Mousavi
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Israel-Iran: How the conflict escalated
On Friday morning, explosions hit Tehran as Israel carried out a major attack on its top army leaders, nuclear sites, and nuclear scientists.
Iran threatened “severe punishment” and quickly retaliated with a wave of missiles.
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Missile aftermath in Israel
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Israeli missile hits warehouse in Iranian city
When questioned about whether Iran could continue fighting Israel, the Iranian ambassador told Yalda Hakim that “it is a matter of principle”.
He said: “This is about self-defence, there is no doubt about it.
“We are a responsible member state of the UN and we do all activities according to our international obligations.
“Any activities are only in the framework of self-defence.”
Image: Damage from an Iranian missile attack to a building in Bat Yam, Israel. Pic: Reuters
Image: Explosions over Jerusalem on Sunday
He added that his country would “do our best to preserve our territorial integrity”, and that “with the help of God”, Iran will “materialise endeavours concretely against our enemy – the Israeli regime”.
Mr Mousavi also told Hakim that Iran’s nuclear activities are “monitored”, and that recent comments by the International Atomic Energy Agency (IAEA) were “politically motivated”.
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Missiles have also been seen over Tel Aviv
The UN nuclear watchdog’s board of governors found Iran was not complying with its nuclear obligations for the first time in 20 years.
Iran said it has “always adhered” to the safeguarding obligations laid down by the watchdog.
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Iranian ambassador reacts to strikes – full interview
Announcing Operation Rising Lion on Friday, Israeli Prime Minister Benjamin Netanyahu claimed Iran had recently taken steps to weaponise enriched uranium, which could be used to make nuclear weapons.
But Mr Mousavi stressed that Iran’s “peaceful activities” at its “nuclear fields” were only for the “generation of electricity, and other peaceful” things.
Iran was due to continue its round of negotiations with the US in Muscat – however, this was cancelled, given recent tensions.
The government is warning people not to travel to Israel under any circumstances, as the country’s missile exchange with Iran shows no sign of abating.
On Friday, the Foreign Office warned against “all but essential travel” to most of Israel.
The areas around Gaza, the West Bank and the Golan Heights were already classed as red zones, with warnings to avoid travel to these areas.
But the government has now updated the warning for the remainder of the country to red.
This puts Israel on the same level as Iran, and the change of advice is also likely to impact travel insurance.
However, with Israel’s airspace closed, it is unlikely many people will be attempting the journey, and Israel’s national airline El Al has announced it is cancelling flights to and from many European cities, as well as Tokyo and Moscow, until 23 June.
The change in travel advice comes after a second night of ballistic missile barrages from Iran following Israel’s attack in the early hours of Friday morning.
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An eight-storey residential building in Tel Aviv was hit by a missile last night.
On Sunday morning, Israel’s health ministry said 12 people had been killed over the past day, taking the total since Friday to 15. It also said 385 people had arrived at hospital with injuries overnight.
Iran has not provided a total number of deaths or overall casualties, but has claimed dozens have been killed.
Iran’s health minister has said most of those injured and killed in Israeli strikes were civilians. According to comments carried by news agency IRNA, he said the majority were women and children.
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The chancellor said UK forces could “potentially” be used to help defend Israel.
The UK government is sending military assets, including fighter jets, to the Middle East.
While the prime minister would not confirm to reporters that UK forces could be used to defend Israel from future Iranian attacks, the chancellor told Sky News earlier that the government is “not ruling anything out”.
Speaking to Sky’s Sunday Morning With Trevor Phillips, Rachel Reeves said sending military assets to the Middle East “does not mean that we are at war”, and emphasised that “we have not been involved in these strikes or this conflict”.
“But we do have important assets in the region,” she continued. “And it is right that we send jets to protect them. And that’s what we’ve done. It’s a precautionary move, and at the same time, we are urging de-escalation.”
Pushed on the question of what the UK would do if Israel asked for support with its operations, the chancellor replied: “I’m not going to rule anything out at this stage. It’s a fast-moving situation, a very volatile situation. But we don’t want to see escalation.”
A helicopter carrying Hindu pilgrims has crashed in India, killing seven people on board.
The accident happened within minutes of the helicopter taking off, officials said, on what should have been a 10-minute flight.
The helicopter was flying to Guptkashi, a prominent Hindu pilgrimage site in the Himalayas, from Kedarnath temple town in the northern Indian state of Uttarakhand.
It comes three days after an Air India flight crashed less than a minute after taking off from Ahmedabad airport in northwestern India, killing at least 270 people.
The helicopter, which was operated by private helicopter service Aryan Aviation, went down in a forested area several miles from the Kedarnath pilgrimage route at around 5.30am local time.
Officials said the crash was believed to have been caused by poor weather conditions.
Authorities say they have launched a search and rescue operation and are expected to review operational protocols for flights in the region.
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The dead include the pilot and pilgrims from the neighbouring state of Uttar Pradesh and western states of Maharashtra and Gujarat, according to officials. The bodies were badly burned in a fire that followed the crash, they said.
Image: Smoke and debris at the site. Pic: Reuters
Tens of thousands of pilgrims visit Kedarnath, which is home to one of the four most sacred Hindu temple shrines, each summer. Many use helicopter services due to the difficult mountainous terrain.
Helicopter mishaps are not uncommon in the region, where sudden weather changes and high-altitude flying conditions can pose risks.
Earlier this month, a helicopter operating in the Kedarnath Valley made an emergency landing shortly after taking off on a highway due to a technical fault. The pilot was injured but all five passengers on board were unharmed.
In May, a helicopter crashed in Uttarkashi district, killing six people, including the pilot. One person survived.